One thing that stands out for me is that there is very little, if any, periphyton.
Which indicates that the systems are oligotrophic (low nutrient).
Possibly, but not likely. In fact, periphyton is present on many oligotrophic systems. Is there epiphytic growth on the plant foliage?Could a healthy submerged plant population not simply replace the presence of periphyton?
That is one form of periphyton. What is the percentage of sand in your sub-strates? Biofilm on sand is of a completely different composition than that any other sub-strate and usually the weakest as to nutrient adsoprtion. It will not usually support any algal growth.Yes. The goldfish and koi constantly graze on it.
The minnows do not.
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